Biography

Carleton College, Northfield, Minnesota: BA (1960)

Stanford University, Stanford, California: PhD (1964)

Postdoctoral Fellow, Stazione Zoologica di Napoli, Naples, italy: 1964-1966

UCSD Faculty (Department of Scripps Institution of Oceanography): 1966-present

Awarded Kowalevsky Medal (jointly with LZ Holland) by the St. Petersburg (Russia) Society of Naturalists: 2014